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Photo: William Vernau

William Vernau- BSc, BVMS, DVSc, PhD
Associate Professor
Pathology, Microbiology & Immunology

3311 Vet Med 3A
Davis, CA 95616

+1 530 754 7162


Research Focus

Hemic neoplasia in dogs and cats, including the characterization of canine T-cell leukemia, a rather poorly characterized constellation of syndromes that often mimic the behavior of their human counterparts and thus have broad comparative biomedical significance. General veterinary clinical pathology with an emphasis on veterinary hematology and cytology, particularly immunophenotyping and immunocytochemistry, and the diagnostic assessment of cerebrospinal fluid.

Specialty Focus

Veterinary Clinical Pathology


1982 BSc , Murdoch University, Perth, Western Australia
1984 BVMS , Murdoch University, Perth, Western Australia
1990 DVSc , University of Guelph, Guelph, Ontario, Canada
2000 PhD , University of California at Davis, Davis, CA, USA

Honors and Awards

2013 Elected to the Teaching Academy of the Consortium of West Region Colleges of Veterinary Medicine
2012 Jerry and Teresa Kaneko Career Development Award in Clinical Pathology
2007 CL Davis Foundation Journal Award
1991 Best graduate student seminar, Department of Pathology, Ontario Veterinary College, University of Guelph.


Diplomate American College of Veterinary Pathology (Clinical Pathology)

Active Research Grants

Principal Investigator, Utility of proliferation indices as prognostic markers for diffuse large B cell lymphoma in dogs, (Principal Investigator), Center for Companion Animal Health

Most Recent 10 Publications

  • Link Icon 2013 Goodnight, AL, Traslavina, RP, Emanuelson, K, Affolter, VK, Gaffney, PM, Vernau, W, Williams, C, Wu, CI, Sturges, BK, Lowenstine, LJ Squamous cell carcinoma of the anal sac in a spotted hyena (Crocuta crocuta). Journal of zoo and wildlife medicine : official publication of the American Association of Zoo Veterinarians,
  • Link Icon 2013 Wiggans K. Tomo, Vernau William, Lappin Michael R., Thomasy Sara M., Maggs David J. Diagnostic utility of aqueocentesis and aqueous humor analysis in dogs and cats with anterior uveitis Veterinary Ophthalmology,
  • Link Icon 2013 O'Brien, D, Moore, PF, Vernau, W, Peauroi, JR, Rebhun, RB, Rodriguez, CO, Skorupski, KA Clinical Characteristics and Outcome in Dogs with Splenic Marginal Zone Lymphoma. Journal of veterinary internal medicine / American College of Veterinary Internal Medicine,
  • Link Icon 2013 Keller, SM, Vernau, W, Hodges, J, Kass, PH, Vilches-Moure, JG, McElliot, V, Moore, PF Hepatosplenic and Hepatocytotropic T-Cell Lymphoma: Two Distinct Types of T-Cell Lymphoma in Dogs. Veterinary Pathology,
  • Link Icon 2013 Kol, A, Christopher, MM, Skorupski, KA, Tokarz, D, Vernau, W B-cell lymphoma with plasmacytoid differentiation, atypical cytoplasmic inclusions, and secondary leukemia in a dog. Veterinary Clinical Pathology,
  • Link Icon 2013 Johnson, LR, Queen, EV, Vernau, W, Sykes, JE, Byrne, BA Microbiologic and Cytologic Assessment of Bronchoalveolar Lavage Fluid from Dogs with Lower Respiratory Tract Infection: 105 Cases (2001-2011). Journal of veterinary internal medicine / American College of Veterinary Internal Medicine,
  • Link Icon 2012 Zwingenberger, AL, Vernau, W, Shi, C, Yan, W, Chen, X, Gordon, IK, Kent, MS Development and characterization of 5 canine B-cell lymphoma cell lines. Leukemia research,
  • Link Icon 2012 Wood, JA, Chung, DJ, Park, SA, Zwingenberger, AL, Reilly, CM, Ly, I, Walker, NJ, Vernau, W, Hayashi, K, Wisner, ER, Cannon, MS, Kass, PH, Cherry, SR, Borjesson, DL, Russell, P, Murphy, CJ Periocular and intra-articular injection of canine adipose-derived mesenchymal stem cells: an in vivo imaging and migration study. Journal of ocular pharmacology and therapeutics : the official journal of the Association for Ocular Pharmacology and Therapeutics,
  • Link Icon 2012 Lane, LV, Allison, RW, Rizzi, TR, Stern, AW, Snider, TA, Moore, PF, Vernau, W Canine intravascular lymphoma with overt leukemia. Veterinary clinical pathology / American Society for Veterinary Clinical Pathology,
  • Link Icon 2012 Mylonakis, ME, Kritsepi-Konstantinou, M, Vernau, W, Valli, VE, Pardali, D, Koutinas, AF Presumptive pure erythroid leukemia in a dog. Journal of veterinary diagnostic investigation : official publication of the American Association of Veterinary Laboratory Diagnosticians, Inc,

Most Recent 5 Book Chapters

2011 Vernau W, Vernau KM, Bolon B: Cerebrospinal Fluid Analysis in Toxicological Neuropathology. Fundamental Neuropathology for Pathologists and Toxicologists: Principles and Techniques, Bolon B, Butt MT, (ed), . 271-283.

2010 Valli T, Vernau W: Classification of Leukemia and Lymphoma, DJ Weiss, K Jane Wardrop, (ed), Schalm's Veterinary Hematology. .

2008 Vernau W, Vernau K, Bailey CS: Cerebrospinal Fluid, Kaneko JJ, Harvey JW, Bruss ML, (ed), Clinical Biochemistry of Domestic Animals, San Diego. 769-819.

2002 Workman HC, Vernau W: Chronic Lymphocytic Leukemia in dogs and cats: the veterinary perspective, , The Veterinary Clinics of North America-Small Animal Practice (Hematology), Philadelphia. 1379-99, viii.

2002 Valli VE, Jacobs RM, Parodi AL, Vernau W, Moore P: WHO International Histological Classification of Tumors of Domestic Animals, , Histological Classification of Hematopoietic Tumors of Domestic Animals. Second Series, Volume VIII, Washington, D.C. .