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Photo: William Vernau

William Vernau- BSc, BVMS, DVSc, PhD
Associate Professor
Pathology, Microbiology & Immunology

3311 Vet Med 3A
Davis, CA 95616

+1 530 754 7162


Research Focus

Hemic neoplasia in dogs, including the characterization of canine T-cell leukemia, a rather poorly charterized constellation of syndromes that often mimic the behavior of their human counterparts and thus have broad comparative biomedical significance.

Specialty Focus

Clinical Pathology


1982 BSc , Murdoch University, Perth, Western Australia
1984 BVMS , Murdoch University, Perth, Western Australia
1990 DVSc , University of Guelph, Guelph, Ontario, Canada
2000 PhD , University of California at Davis, Davis, CA, USA

Honors and Awards

2007 CL Davis Foundation Journal Award


Diplomate American College of Veterinary Pathology (Clinical Pathology)

Active Research Grants

Co-Investigator, Comparison of cyclosporine and prednisone in the treatment of primary immune-mediated polyarthritis in dog, Dr. Jane Sykes (Principal Investigator), Center for Companion Animal Health

Principal Investigator, Investigation of the effects of sample type and collection tube type on biochemical analytes in horses, (Principal Investigator), Center for Equine Health

Most Recent 10 Publications

  • Link Icon 2011 Gilroy C, Forzán M, Drew A, Vernau W Eosinophilia in a cat with acute leukemia Canadian Veterinary Journal,
  • Link Icon 2011 Osofsky A, Hawkins MG, Foreman O, Kent MS, Vernau W, Lowenstine LJ T-Cell Chronic Lymphocytic Leukemia in a Double Yellow- Headed Amazon Parrot (Amazona ochrocephala oratrix) Journal of Avian Medicine and Surgery,
  • Link Icon 2011 Mylonakis ME, Soubasis N, Kritsepi-Konstantinou M, Vernau W, Theodorou K, Tentoma L, Koutinas AF Presumptive histiocytic neoplasm with unusual immunophenotype in a cat Comp Clin Pathol DOI 10.1007/s00580-011-1304-6,
  • Link Icon 2011 Cruz Cardona JA, Milner R, Alleman AR, Williams C, Vernau W, Breen M, Tompkins M BCR-ABL translocation in a dog with chronic monocytic leukemia Vet Clin Pathol,
  • Link Icon 2011 Valli VE, San Myint M, Barthel A, Bienzle D, Caswell J, Colbatzky F, Durham A, Ehrhart EJ, Johnson Y, Jones C, Kiupel M, Labelle P, Lester S, Miller M, Moore P, Moroff S, Roccabianca P, Ramos-Vara J, Ross A, Scase T, Tvedten H, Vernau W Classification of canine malignant lymphomas according to the World Health Organization criteria Vet Pathol,
  • Link Icon 2011 Juopperi TA, Bienzle D, Bernreuter DC, Vernau W, Thrall MA, McManus PM Prognostic markers for myeloid neoplasms: a comparative review of the literature and goals for future investigation Vet Pathol,
  • Link Icon 2011 Webster JD, Dennis MM, Dervisis N, Heller J, Bacon NJ, Bergman PJ, Bienzle D, Cassali G, Castagnaro M, Cullen J, Esplin DG, Peña L, Goldschmidt MH, Hahn KA, Henry CJ, Hellmén E, Kamstock D, Kirpensteijn J, Kitchell BE, Amorim RL, Lenz SD, Lipscomb TP, McEntee M, McGill LD, McKnight CA, McManus PM, Moore AS, Moore PF, Moroff SD, Nakayama H, Northrup NC, Sarli G, Scase T, Sorenmo K, Schulman FY, Shoieb AM, Smedley RC, Spangler WL, Teske E, Thamm DH, Valli VE, Vernau W, von Euler H, Withrow SJ, Weisbrode SE, Yager J, Kiupel M Recommended guidelines for the conduct and evaluation of prognostic studies in veterinary oncology Vet Pathol,
  • Link Icon 2011 Johnson LR, Vernau W Bronchoscopic findings in 48 cats with spontaneous lower respiratory tract disease (2002-2009) J Vet Intern Med,
  • Link Icon 2011 Bienzle D, Vernau W The diagnostic assessment of canine lymphoma: implications for treatment Clinics in Laboratory Medicine,
  • Link Icon 2011 Kennedy DR, Hartnett BJ, Kennedy JS, Vernau W, Moore PF, O'Malley T, Burkly LC, Henthorn PS, Felsburg PJ Ex vivo ?-retroviral gene therapy of dogs with X-linked severe combined immunodeficiency and the development of a thymic T cell lymphoma Vet Immunol Immunopathol,

Most Recent 5 Book Chapters

2011 Vernau W, Vernau KM, Bolon B: Cerebrospinal Fluid Analysis in Toxicological Neuropathology. Fundamental Neuropathology for Pathologists and Toxicologists: Principles and Techniques, Bolon B, Butt MT, (ed), . 271-283.

2010 Valli T, Vernau W: Classification of Leukemia and Lymphoma, DJ Weiss, K Jane Wardrop, (ed), Schalm's Veterinary Hematology. .

2008 Vernau W, Vernau K, Bailey CS: Cerebrospinal Fluid, Kaneko JJ, Harvey JW, Bruss ML, (ed), Clinical Biochemistry of Domestic Animals, San Diego. 769-819.

2002 Workman HC, Vernau W: Chronic Lymphocytic Leukemia in dogs and cats: the veterinary perspective, , The Veterinary Clinics of North America-Small Animal Practice (Hematology), Philadelphia. 1379-99, viii.

2002 Valli VE, Jacobs RM, Parodi AL, Vernau W, Moore P: WHO International Histological Classification of Tumors of Domestic Animals, , Histological Classification of Hematopoietic Tumors of Domestic Animals. Second Series, Volume VIII, Washington, D.C. .