Walter Boyce

Walter Boyce has been Executive Director of the UC Davis Wildlife Health Center since 1998, and has been a professor and wildlife veterinarian on the faculty of the UC Davis School of Veterinary Medicine since 1988. His educational background includes zoology (BS, Auburn University 1978), veterinary medicine (DVM, Auburn University 1981), and parasitology (MS, University of Florida 1985; PhD, Purdue University 1988).

He teaches parasitology and wildlife health and disease to veterinary students at UC Davis, and he supervises graduate students in the following graduate groups; Ecology, Epidemiology, Comparative Pathology.

Walter is interested in a wide range of issues and has published over 80 scientific papers on species ranging from Africanized bees to bighorn sheep, and in disciplines ranging from ecology to molecular genetics. In addition to mountain lions, he currently oversees projects on West Nile Virus and Avian Influenza.

PUBLICATIONS

BY DATE; click HERE for PUBLICATIONS BY TOPIC. Clicking on a link within publications will open a pdf version of the article.

2005; Bauer, J.W., K.A. Logan, L.L. Sweanor, and W.M. Boyce. Scavenging behavior in puma. Southwestern Naturalist. 50:466-471. pdf

2005; Boyce, W.M. and M. Weisenberger. The rise and fall of psoroptic scabies in bighorn sheep in the San Andres Mountains, New Mexico. Journal of Wildlife Diseases. 41:525—531. pdf

2005; Teglas, M.B., May, B.P. Crosbie, P.R., Stevens, M.R., and W.M. Boyce. Genetic structure of the tick Ornithodoros coriaceus (Acari: Argasidae) in California, Nevada and Oregon. Journal of Medical Entomology. 42: 247-253. pdf

2005; Padgett, K.A, and W.M. Boyce. Ants as first intermediate host of Mesocestoides on San Miguel Island, USA. Journal of Helminthology. 79:67-73. pdf

2005; Padgett, K.A, S.A. Nadler, L. Munson, Sacks, B. and W.M. Boyce. Systematics of Mesocestoides (Cestoda: Mesocestoididae):  Evaluation of molecular and morphological variation among isolates. Journal of Parasitology. 91:1435-1443. pdf

2004; Padgett, K.A, and W.M. Boyce. Life history studies on two molecular strains of Mesocestoides (Cestoda: Mesocestoididae): identification of sylvatic hosts and infectivity of immature life stages. Journal of Parasitology. 90:108-113. pdf

2004; Whittaker, D.G., S.D. Ostermann, and W.M. Boyce. Genetic variability of reintroduced California bighorn sheep in Oregon. Journal of Wildlife Management. 68:850-859.

2003; Ernest, H.B., W.M. Boyce, V.C. Bleich, B. May, S.J. Stiver and S.G. Torres. Genetic structure of mountain lion (Puma concolor) populations in California. Conservation Genetics. pdf

2003; Boyce, W.M., C.S. O’Brien and E.S. Rubin. Responses of bighorn sheep (Ovis canadensis) to feral honey bees (Apis mellifera) at water. Southwestern Naturalist. 48:81-84.

2003; McLaughlin, B.E., C.D. DeMaula, W.C. Wilson, W.M. Boyce, and N.J. MacLachlan. Replication of bluetongue virus and epizootic hemorrhagic disease virus in pulmonary artery endothelial cells obtained from cattle, sheep, and deer. American Journal of Veterinary Research. 64:860-865. pdf

2002; Ernest, H.B., E.S. Rubin and W.M. Boyce. Fecal DNA analysis and risk assessment of mountain lion predation of bighorn sheep. Journal of Wildlife Management 66(1):75-85. pdf

2002; Rubin, E.S., W.M. Boyce, C.J. Sterner and S.G. Torres. Bighorn sheep habitat use and selection near an urban environment. Biological Conservation 104:251-263. pdf

2002; Ziccardi, M.H., J.K. Mazet, I.A. Gardner, W.M. Boyce and M.S. Denison. Validation of a cell culture bioassay for detection of petroleum exposure in mink (Mustela vison) as a model for sea otter (Enhydra lutris). American Journal of Veterinary Research 63(7):963-968. pdf

2002; Rubin, E.S., Boyce, W.M., and E.P. Caswell-Chen.  Modeling demographic processes in an endangered population of bighorn sheep. Journal of Wildlife Management 66(3):796-810. pdf

2002; Boyce, W.M., E.S. Rubin and C.S. O’Brien. A scientific note on the distribution of Africanized honey bees and Varroa destructor in feral honey bee populations in California. Apidologie 33:581-582.

2001; Rodwell, T.C., I.J. Whyte and W.M. Boyce. Evaluation of population effects of bovine tuberculosis in free-ranging African buffalo (Syncerus caffer). Journal of Mammalogy 82(1)231-238. pdf

2001; Gilardi, K.V.K., J.D. Gilardi, A. Frank, M.L. Goff and W.M. Boyce. Epidermoptid mange in Laysan albatross fledglings in Hawaii. Journal of Wildlife Diseases 37(1):185-188. pdf

2001; Rodwell, T.C., N.P. Kriek, R.C. Bengis, I.J. Whyte, P.C. Viljoen, V. de Vos, and W.M. Boyce. Prevalence of bovine tuberculosis in African buffalo at Kruger National Park. Journal of Wildlife Diseases 37(2):258-264. pdf

2001; Gutierrez-Espeleta, G.A., P.W. Hedrick, S.T. Kalinowski, D. Garrigan and W.M. Boyce. Is the decline of desert bighorn sheep from infectious disease the result of low MHC variation? Heredity 86:439-450. pdf

2000; Borjesson, D.L., M.C. Christopher and W.M. Boyce. Biochemical and hematologic reference intervals for free-ranging desert bighorn sheep. Journal of Wildlife Diseases 36:294-300. pdf

2000; Crosbie, P.R., S.A. Nadler, E.G. Platzer, C. Kerner, J. Mariaux and W.M. Boyce. Molecular systematics of Mesocestoides spp. (Cestoda: Mesocestoididae) from domestic dogs (Canis familiaris) and coyotes (Canis latrans). Journal of Parasitology 86:350-357. pdf

2000; Ernest, H.B., M.C.T. Penedo, B.P. May, M. Syvanen and W.M. Boyce. Molecular tracking of mountain lions in the Yosemite Valley region of California: genetic analysis using microsatellites and fecal DNA. Molecular Ecology 9:433-442. pdf

2000; Sweitzer, RA, D. Van Vuren, I.A. Gardner, W.M. Boyce and J.D. Waithman. Estimating sizes of wild pig populations in the North and Central Coast regions of California. Journal of Wildlife Management 64:531-543.

2000; Chang, C., B.B. Chomel, R.W. Kasten, R. Heller, K.M. Kocan, H. Ueno, K. Yamamoto, V.C. Bleich, B.M. Pierce, B.J. Gonzales, P.K. Swift, W.M. Boyce, S.S. Yang, H. Boulouis and Y. Piemont. Bartonella spp. isolated from wild and domestic ruminants in North America. Emerging Infectious Diseases 6:306-311.

2000; Rubin, E.S., W.M. Boyce and V.C. Bleich. Reproductive strategies of desert bighorn sheep. Journal of Mammalogy 81(3):769-786. pdf

2000; Hayes, C.L., E.S. Rubin, M.C. Jorgensen, R.A. Botta and W.M. Boyce.
Mountain lion predation on bighorn sheep in the Peninsular Ranges, California. Journal of Wildlife Management 64:954-959.

2000; Ramey, R.R., S.T. Kelley, W.M. Boyce and B.D. Farrell. Phylogeny and host specificity of psoroptic mange mites (Acarina:Psoroptidae) as indicated by ITS sequence data. Journal of Medical Entomology 37(6):791-796. pdf

2000; Mazet, J.K., I.A. Gardner, D. A. Jessup, L.J. Lowenstine and W.M. Boyce.  Evaluation of changes in hematologic and clinical biochemical values after exposure to petroleum products in mink (Mustela vison) as a model for assessment of sea otters (Enhydra lutris) American Journal of Veterinary Research 61(10):1197-1203. pdf

2000; Gutiérrez-Espeleta, G.A., S.T. Kalinowski, W.M. Boyce and P.W. Hedrick. Genetic variation and population structure in desert bighorn sheep: implications for conservation. Conservation Genetics 1:3-15. pdf

2000; Boyce, W.M., L.W. Woods, K. Keel, N.J. MacLachlan, C.O. Porter and H. D.
Lehmkuhl. An epizootic of adenovirus-induced hemorrhagic disease in captive black-tailed deer (Odocoileus hemionus). Journal of Zoo and Wildlife Medicine 31(3):370-373. pdf

1999; Boyce, W.M., R.R. Ramey, T.C. Rodwell, E.S. Rubin and R.S. Singer. Population subdivision among bighorn sheep (Ovis canadensis) ewes revealed by mitochondrial DNA analysis. Molecular Ecology 8:99-106.

1999; Waithman, J.D., R.A. Sweitzer, D.Van Vuren, J.D. Drew, A.J. Brinkhaus, I.A. Gardner and W.M. Boyce. Range expansion, population sizes, and management of wild pigs in California. Journal of Wildlife Management 63:298-308.

1999; Stiller, D., P.R. Crosbie, W.M. Boyce and W.L. Goff.  Dermacentor hunteri (Acari: Ixodidae): an experimental vector of Anaplasma marginale and A. ovis (Rickettsiales: Anaplasmataceae) to calves and sheep. Journal of Medical Entomology 36:321-324

1999; Boyce, W., A. Fisher, H. Provencio, E. Rominger, J. Thilstead and M. Ahlm. Elaeophorosis in bighorn sheep in New Mexico. Journal of Wildlife Diseases 35:786-789. pdf

1998; Singer, R.S., W.M. Boyce, I.A. Gardner, W. O. Johnson and A. Fisher. Evaluation of bluetongue virus diagnostic tests in free-ranging bighorn sheep. Preventive Veterinary Medicine 12:1-18. pdf

1998; Crosbie, P.R., W.M. Boyce and T.C. Rodwell. DNA sequence variation in Dermacentor hunteri and estimated phylogenies of Dermacentor spp. (Acari: Ixodidae) in the New World. Journal of Medical Entomology 35:277-288.

1998; Crosbie, P.R. and W.M. Boyce. Dermacentor hunteri (Acari: Ixodidae): Seasonal variation in questing adults and on-host juvenile stages, and host associations and feeding behavior of larvae and nymphs. Journal of Medical Entomology 35:1034-1043.

1998; Crosbie, P.R., W.M. Boyce, E.G. Platzer, S.A. Nadler and C. Kerner. Diagnostic procedures and treatment of eleven dogs with peritoneal infections caused by Mesocestoides spp. Journal of the American Veterinary Medical Association 213:1578-1583.

1998; Rubin, E.S., W.M. Boyce, M.C. Jorgensen, S.G. Torres, C.L. Hayes, C.S. O’Brien and D.A. Jessup. Distribution and abundance of bighorn sheep in the Peninsular Ranges, California. Wildlife Society Bulletin 26:539-551.

1997; Singer, R.S., D.A. Jessup, I.A. Gardner and W.M. Boyce. Pathogen exposure patterns among sympatric populations of bighorn sheep (Ovis canadensis), mule deer (Odocoileus hemionus), and cattle. Journal of Wildlife Diseases 33:377-382. pdf

1997; Boyce, W.M., P.W. Hedrick, N.E. Muggli-Cockett, S. Kalinowski, C.T. Penedo and R.R. Ramey. Genetic variation of major histocompatibility complex and microsatellite loci: a comparison in bighorn sheep. Genetics 145:421-433. pdf

1997; Crosbie, P.R., W.L. Goff, D. Stiller, D.A. Jessup and W.M. Boyce. The distribution of Dermacentor hunteri and Anaplasma sp. in bighorn sheep. Journal of Parasitology 83:31-37. pdf

1997; Sweitzer, R.A., G.S. Ghneim, I.A. Gardner, D.Van Vuren, B.J. Gonzales and W.M. Boyce. Immobilization and physiological parameters associated with chemical restraint of wild pigs with Telazol and xylazine hydrochloride. Journal of Wildlife Diseases 33:198-205. pdf

1997; Atwill, E.R., R.A. Sweitzer, M. G. Pereira, I. A. Gardner, D. Van Vuren and W.M. Boyce. Prevalence of and associated risk factors for shedding Cryptosporidium parvum and Giardia duodenalis within feral pig populations in California. Applied and Environmental Microbiology 63:3946-3949. pdf

1997; Sweitzer, R.A, B.J. Gonzales, I.A. Gardner, D.Van Vuren, W.M. Boyce and J.D. Waithman. A modified panel trap and immobilization technique for capturing multiple wild pigs. Wildlife Society Bulletin 25:699-705.

1996; Ziccardi, M., W.M. Boyce and I. Gardner. Seroprevalence of Psoroptes sp. mites in free-ranging elk (Cervus elaphus) as determined by kinetic ELISA. Journal of Wildlife Diseases 32:51-56. pdf

1996; Borjesson, D., Boyce, W.M., I.A. Gardner, J. DeForge and B. Lasley. Pregnancy detection in bighorn sheep using a fecal-based enzyme immunoassay. Journal of Wildlife Diseases 32:67-74. pdf

1996; Reese, N.E., W.M. Boyce, I.A. Gardner and D.M. Nelson. Fixation affects morphometric characters of Psoroptes cuniculi mites (Acari:Psoroptidae). Journal of Medical Entomology 33:835-838.

1996; Boyce, W.M. and R.L. Zarnke. Antibody responses to Psoroptes sp. mites in Dall sheep (Ovis dalli). Journal of Wildife Diseases 32:711-713 pdf

1996; Lee, G.Y., W.M, Boyce and K. Orr. Diagnosis and treatment of lungworm (Filariopsis arator; Metastrongyloidea: Filaroididae) infection in white-faced capuchins (Cebus capucinus). Journal of Zoo and Wildlife Medicine 27:197-200.

1996; Gardner, I.A., S. Hietala and W.M. Boyce. Validity of using serologic tests for diagnosis of diseases in wild animals. Reviews of Science and Technology Office of International Epizootics 15:323-335.

1995; Kjemtrup, A., J. Thomford, I. Gardner, P.A. Conrad, D.A. Jessup and W.M. Boyce. Seroprevalence of two Babesia spp. isolates in selected bighorn sheep (Ovis canadensis) and mule deer (Odocoileus hemionus) populations in California. Journal of Wildlife Diseases 31:467-471.

1994; Elliott, L.F., W.M. Boyce, R.K. Clark and D.A. Jessup. Geographic analysis of pathogen exposure in bighorn sheep (Ovis canadensis). Journal of Wildlife Diseases 30:315-318. pdf

1994; Mikolon, A.B., W.M. Boyce, J.L. Allen, I.A. Gardner and L.F. Elliott. Epidemiology and control of nematode parasites in a collection of captive exotic ungulates. Journal of Zoo and Wildlife Medicine 25:500-510.

1993; Clark, R.K., W.M. Boyce, D.A. Jessup and L.F. Elliott. Survey of pathogen exposure among population clusters of in bighorn sheep (Ovis canadensis) in California. Journal of Zoo and Wildlife Medicine 24:48-53.

1993; Thomford, J.W., P.A. Conrad, W.M. Boyce, P.J. Holman and D.A. Jessup. Isolation and in vitro cultivation of Babesia parasites from free-ranging desert bighorn sheep (Ovis canadensis nelsoni) and mule deer (Odocoileus hemionus) in California. Journal of Parasitology 79:77-84. pdf

1993; Boyce, W.M. and A. Kjemtrup. Wildlife health and the urban-wildlife interface. Journal of the American Veterinary Medical Association 202:1082-1084.

1993; Goff, W.L., D.A. Jessup, K.A. Waldrup, J.W. Thomford, P.A. Conrad, W.M. Boyce, J.R. Gorham and G.A. Wagner. The isolation and partial characterization of a Babesia sp. from desert bighorn sheep (Ovis canadensis nelsoni). The Journal of Eucaryotic Microbiology 40:237-243.

1993; Goff, W., D. Stiller, D. Jessup, P. Msolla, W.M. Boyce and W. Foreyt. Characterization of an Anaplasma ovis isolate from desert bighorn sheep in southern California. Journal of Wildlife Diseases 29:540-546.

1993; Jessup, D.A., W.L. Goff, D. Stiller, M.N. Oliver, V.C. Bleich and W.M. Boyce. A retrospective serologic survey for Anaplasma spp. infection in three bighorn sheep (Ovis canadensis) populations in California Journal of Wildlife Diseases. 29:547-554. pdf

1993; Murphy, F.A., D.L. Brooks, W.M. Boyce, W. Lasley, L. Lowenstine, J.W. Glosser, B.I. Osburn, M.E. Fowler, D.A. Jessup, A. Ardans and W.P. Heuschele. An international system for the prevention and control of infectious diseases in free-ranging and captive wild animals. Journal Zoo and Wildlife Medicine 24:365-373.

1993; Goldberg, M.A., K.R. Kazacos, W.M. Boyce, E. Abi and B. Katz. California DUSN: Morphometric, serologic and epidemiologic support for Baylisascaris as a causative agent. Ophthalmology 100:1695-1701.

1992; Boyce, W.M., T. Yuill, J. Homan and D.A. Jessup. A role for veterinarians in wildlife health and conservation biology. Journal of the American Veterinary Medical Association 200:435-437.

1992; Foreyt, W.J., L.G. Rickard and W.M. Boyce. Psoroptes sp. in two llamas in Washington. Journal of Parasitology 78:153-155.

1992; Boyce, W.M., R.N. Brown, B.C. Zingg, R.B. Lefebvre, and R.S. Lane. First isolation of Borrelia burgdorferi from southern California. Journal of Medical Entomology 29:496-500.

1992; Boyce, W.M., J.A. Miller, D.A. Jessup and R.K. Clark. Use of ivermectin implants for the treatment of psoroptic scabies in free-ranging bighorn sheep. Journal of Zoo and Wildlife Medicine 23:211-213.

1992; Mazet, J.A.K., W.M. Boyce, J. Mellies, R.K. Clark and D.A. Jessup. Exposure to Psoroptes sp. mites is common among bighorn sheep (Ovis canadensis) populations in California. Journal of Wildlife Diseases 28:542-547. pdf

1992; Elliott, L.F. and W.M. Boyce. Implications of captive breeding programs for the conservation of bighorn sheep. Transactions of the Desert Bighorn Council 36:54-57.

1991; Boyce, W.M. and E.A. Kazacos. Histopathology of nymphal pentastomid infections (Sebekia mississippiensis) in paratenic hosts. Journal of Parasitology 77:104-110.

1991; Boyce, W.M., D.A. Jessup and R.K. Clark. Serodiagnostic antibody responses to Psoroptes sp. infestations in bighorn sheep. Journal of Wildlife Diseases 27:10-15.

1991; Boyce, W.M., J.L. Allen, C. Himmelwright, L. Elliott, A. Mikolon, J. Mazet and I. Gardner. Implementation and evaluation of a strategic parasite control program for captive exotic ungulates. Journal of the American Veterinary Medical Association 198:1972-1976.

1991; Boyce, W.M. and R.N. Brown. Antigenic characterization of Psoroptes spp. (Acari: Psoroptidae) mites from different hosts. Journal of Parasitology 77:675-679.

1991; Boyce, W.M., J.A.K. Mazet, J. Mellies, I. Gardner, R.K. Clark and D.A. Jessup. Kinetic ELISA for detection of antibodies to Psoroptes sp. (Acari: Psoroptidae) in bighorn sheep (Ovis canadensis). Journal of Parasitology 77:692-696.

1990; Boyce, W.M., C.H. Courtney and P.E. Loggins. Seasonal transmission of Fasciola hepatica in north-central Florida. (U.S.A.). International Journal for Parasitology 20:695-696.

1990; Boyce, W.M., L. Elliott, R.K. Clark and D.A. Jessup. Morphometric analysis of Psoroptes spp. mites from bighorn sheep, mule deer, rabbits and cattle. Journal of Parasitology 76:823-828.

1989; Boyce, W.M., D.J. Asai, J.K. Wilder and K.R. Kazacos. Physicochemical characterization and monoclonal and polyclonal antibody recognition of Baylisascaris procyonis larval excretory - secretory antigens. Journal of Parasitology 75:540-548.

1989; Kazacos, K.R. and W.M. Boyce. Baylisascaris larva migrans. Journal of the American Veterinary Medical Association 195:894-903.

1988; Boyce, W.M., B.A. Branstetter and K.R. Kazacos. In vitro culture of Baylisascaris procyonis and initial analysis of larval excretory-secretory antigens. Proceedings of the Helminthological Society of Washington 55(1):15-18.

1988; Boyce, W.M., B.A. Branstetter and K.R. Kazacos. Comparative analysis of larval excretory-secretory antigens of Baylisascaris procyonis, Toxocara canis and Ascaris suum by western blotting and enzyme immunoassay. International Journal for Parasitology 18(1):109-113.

1987; Boyce, W.M., C.H. Courtney, S.R. Wing and E.W. Kurose.  In vitro maintenance of the pentastome Sebekia mississippiensis. Proceedings of the Helminthological Society of Washington 54(2):265-266.

1987; Boyce, W.M., C.H. Courtney and P.E. Loggins. Resistance to experimental infection with Fasciola hepatica in exotic and domestic breeds of sheep. International Journal for Parasitology 17(7):1233-1237.

1987; Boyce, W.M., E.A. Kazacos, K.R. Kazacos and J.A. Engelhardt. Pathology of pentastomid infections (Sebekia mississippiensis) in fish. Journal of Wildlife Diseases 23(4):689-692.

1986; Boyce, W.M., C.H. Courtney and M.B. Thibideau.  Egg production of Pseudosuccinea columella and Fossaria cubensis (Lymnaeidae) in laboratory culture.  Journal of Parasitology 72(1):184.

1986; Boyce, W.M., C.H. Courtney and M.B. Thibideau.  Heterologous resistance to Fasciola hepatica conferred upon rats by passive transfer of serum from different breeds of sheep.  Veterinary Parasitology 22:259-266.

1985; Boyce, W.M. The prevalence of Sebekia mississippiensis (Pentastomida) in American alligators (Alligator mississippiensis) in North Florida and experimental infection of paratenic hosts.  Proceedings of the Helminthological Society of Washington 52(2):278-282.

1985; Allen, J.L., E.R. Jacobson, J.W. Harvey and W.M. Boyce.  Hematologic and serum chemical values for young African elephants (Loxodonta africana) with variations for sex and age.  Journal of Zoo Animal Medicine 16:98-101.

1985; Allen, J.L., G.V. Kollias, E.C. Greiner and W.M. Boyce.  Subcutaneous filariasis (Pelecitus sp.) in a yellow-collared Macaw (Ara auricollis)  Avian Diseases. 29(3):891-894.

1984; Boyce, W.M., P. Cardeilhac, T. Lane, C. Buergelt and M. King.  Sebekiosis in captive alligator hatchlings.  Journal of the American Veterinary Medical Association 185(11):1419-1420.

1984; Boyce, W.M., G. Kollias, C.H. Courtney, J.L. Allen and E. Chalmers.  Efficacy of ivermectin against gastrointestinal nematodes in dromedary camels.  Journal of the American Veterinary Medical Association 185(11):1307-1308.

Publications by topic

Lions

2005; Bauer, J.W., K.A. Logan, L.L. Sweanor, and W.M. Boyce. Scavenging behavior in puma. Southwestern Naturalist. 50:466-471. pdf

2003; Ernest, H.B., W.M. Boyce, V.C. Bleich, B. May, S.J. Stiver and S.G. Torres. Genetic structure of mountain lion (Puma concolor) populations in California. Conservation Genetics. pdf

2002; Ernest, H.B., E.S. Rubin and W.M. Boyce. Fecal DNA analysis and risk assessment of mountain lion predation of bighorn sheep. Journal of Wildlife Management 66(1):75-85. pdf

2000; Ernest, H.B., M.C.T. Penedo, B.P. May, M. Syvanen and W.M. Boyce. Molecular tracking of mountain lions in the Yosemite Valley region of California: genetic analysis using microsatellites and fecal DNA. Molecular Ecology 9:433-442. pdf

2000; Hayes, C.L., E.S. Rubin, M.C. Jorgensen, R.A. Botta and W.M. Boyce.
Mountain lion predation on bighorn sheep in the Peninsular Ranges, California. Journal of Wildlife Management 64:954-959.

Bighorn Sheep

2005; Boyce, W.M. and M. Weisenberger. The rise and fall of psoroptic scabies in bighorn sheep in the San Andres Mountains, New Mexico. Journal of Wildlife Diseases. 41:525-531. pdf

2004; Whittaker, D.G., S.D. Ostermann, and W.M. Boyce. Genetic variability of reintroduced California bighorn sheep in Oregon. Journal of Wildlife Management. 68:850-859.

2003; Boyce, W.M., C.S. O’Brien and E.S. Rubin. Responses of bighorn sheep (Ovis canadensis) to feral honey bees (Apis mellifera) at water. Southwestern Naturalist. 48:81-84.

2002; Ernest, H.B., E.S. Rubin and W.M. Boyce. Fecal DNA analysis and risk assessment of mountain lion predation of bighorn sheep. Journal of Wildlife Management 66(1):75-85. pdf

2002; Rubin, E.S., W.M. Boyce, C.J. Sterner and S.G. Torres. Bighorn sheep habitat use and selection near an urban environment. Biological Conservation 104:251-263. pdf

2002; Rubin, E.S., Boyce, W.M., and E.P. Caswell-Chen.  Modeling demographic processes in an endangered population of bighorn sheep. Journal of Wildlife Management 66(3):796-810.

2001; Gutierrez-Espeleta, G.A., P.W. Hedrick, S.T. Kalinowski, D. Garrigan and W.M. Boyce. Is the decline of desert bighorn sheep from infectious disease the result of low MHC variation? Heredity 86:439-450. pdf

2000; Borjesson, D.L., M.C. Christopher and W.M. Boyce. Biochemical and hematologic reference intervals for free-ranging desert bighorn sheep. Journal of Wildlife Diseases 36:294-300. pdf

2000; Chang, C., B.B. Chomel, R.W. Kasten, R. Heller, K.M. Kocan, H. Ueno, K. Yamamoto, V.C. Bleich, B.M. Pierce, B.J. Gonzales, P.K. Swift, W.M. Boyce, S.S. Yang, H. Boulouis and Y. Piemont. Bartonella spp. isolated from wild and domestic ruminants in North America. Emerging Infectious Diseases 6:306-311.

2000; Rubin, E.S., W.M. Boyce and V.C. Bleich. Reproductive strategies of desert bighorn sheep. Journal of Mammalogy 81(3):769-786. pdf

2000; Hayes, C.L., E.S. Rubin, M.C. Jorgensen, R.A. Botta and W.M. Boyce.
Mountain lion predation on bighorn sheep in the Peninsular Ranges, California. Journal of Wildlife Management 64:954-959.

2000; Ramey, R.R., S.T. Kelley, W.M. Boyce and B.D. Farrell. Phylogeny and host specificity of psoroptic mange mites (Acarina:Psoroptidae) as indicated by ITS sequence data. Journal of Medical Entomology 37(6):791-796. pdf

2000; Gutiérrez-Espeleta, G.A., S.T. Kalinowski, W.M. Boyce and P.W. Hedrick. Genetic variation and population structure in desert bighorn sheep: implications for conservation. Conservation Genetics 1:3-15. pdf

1999; Boyce, W.M., R.R. Ramey, T.C. Rodwell, E.S. Rubin and R.S. Singer. Population subdivision among bighorn sheep (Ovis canadensis) ewes revealed by mitochondrial DNA analysis. Molecular Ecology 8:99-106. pdf

1999; Waithman, J.D., R.A. Sweitzer, D.Van Vuren, J.D. Drew, A.J. Brinkhaus, I.A. Gardner and W.M. Boyce. Range expansion, population sizes, and management of wild pigs in California. Journal of Wildlife Management 63:298-308.

1999; Boyce, W., A. Fisher, H. Provencio, E. Rominger, J. Thilstead and M. Ahlm. Elaeophorosis in bighorn sheep in New Mexico. Journal of Wildlife Diseases 35:786-789.

1998; Singer, R.S., W.M. Boyce, I.A. Gardner, W. O. Johnson and A. Fisher. Evaluation of bluetongue virus diagnostic tests in free-ranging bighorn sheep. Preventive Veterinary Medicine 12:1-18.

1998; Rubin, E.S., W.M. Boyce, M.C. Jorgensen, S.G. Torres, C.L. Hayes, C.S. O’Brien and D.A. Jessup. Distribution and abundance of bighorn sheep in the Peninsular Ranges, California. Wildlife Society Bulletin 26:539-551.

1997; Singer, R.S., D.A. Jessup, I.A. Gardner and W.M. Boyce. Pathogen exposure patterns among sympatric populations of bighorn sheep (Ovis canadensis), mule deer (Odocoileus hemionus), and cattle. Journal of Wildlife Diseases 33:377-382. pdf

1997; Boyce, W.M., P.W. Hedrick, N.E. Muggli-Cockett, S. Kalinowski, C.T. Penedo and R.R. Ramey. Genetic variation of major histocompatibility complex and microsatellite loci: a comparison in bighorn sheep. Genetics 145:421-433.

1997; Crosbie, P.R., W.L. Goff, D. Stiller, D.A. Jessup and W.M. Boyce. The distribution of Dermacentor hunteri and Anaplasma sp. in bighorn sheep. Journal of Parasitology 83:31-37. pdf

1996; Borjesson, D., Boyce, W.M., I.A. Gardner, J. DeForge and B. Lasley. Pregnancy detection in bighorn sheep using a fecal-based enzyme immunoassay. Journal of Wildlife Diseases 32:67-74. pdf

1996; Boyce, W.M. and R.L. Zarnke. Antibody responses to Psoroptes sp. mites in Dall sheep (Ovis dalli). Journal of Wildife Diseases 32:711-713 pdf

1995; Kjemtrup, A., J. Thomford, I. Gardner, P.A. Conrad, D.A. Jessup and W.M. Boyce. Seroprevalence of two Babesia spp. isolates in selected bighorn sheep (Ovis canadensis) and mule deer (Odocoileus hemionus) populations in California. Journal of Wildlife Diseases 31:467-471.

1994; Elliott, L.F., W.M. Boyce, R.K. Clark and D.A. Jessup. Geographicanalysis of pathogen exposure in bighorn sheep (Ovis canadensis). Journal of Wildlife Diseases 30:315-318.

1993; Clark, R.K., W.M. Boyce, D.A. Jessup and L.F. Elliott. Survey of pathogen exposure among population clusters of in bighorn sheep (Ovis canadensis) in California. Journal of Zoo and Wildlife Medicine 24:48-53.

1993; Thomford, J.W., P.A. Conrad, W.M. Boyce, P.J. Holman and D.A. Jessup. Isolation and in vitro cultivation of Babesia parasites from free-ranging desert bighorn sheep (Ovis canadensis nelsoni) and mule deer (Odocoileus hemionus) in California. Journal of Parasitology 79:77-84. pdf

1993; Goff, W.L., D.A. Jessup, K.A. Waldrup, J.W. Thomford, P.A. Conrad, W.M. Boyce, J.R. Gorham and G.A. Wagner. The isolation and partial characterization of a Babesia sp. from desert bighorn sheep (Ovis canadensis nelsoni). The Journal of Eucaryotic Microbiology 40:237-243.

1993; Goff, W., D. Stiller, D. Jessup, P. Msolla, W.M. Boyce and W. Foreyt. Characterization of an Anaplasma ovis isolate from desert bighorn sheep in southern California. Journal of Wildlife Diseases 29:540-546.

1993; Jessup, D.A., W.L. Goff, D. Stiller, M.N. Oliver, V.C. Bleich and W.M. Boyce. A retrospective serologic survey for Anaplasma spp. infection in three bighorn sheep (Ovis canadensis) populations in California Journal of Wildlife Diseases. 29:547-554. pdf

1992; Boyce, W.M., J.A. Miller, D.A. Jessup and R.K. Clark. Use of ivermectin implants for the treatment of psoroptic scabies in free-ranging bighorn sheep. Journal of Zoo and Wildlife Medicine 23:211-213.

1992; Mazet, J.A.K., W.M. Boyce, J. Mellies, R.K. Clark and D.A. Jessup. Exposure to Psoroptes sp. mites is common among bighorn sheep (Ovis canadensis) populations in California. Journal of Wildlife Diseases 28:542-547.

1992; Elliott, L.F. and W.M. Boyce. Implications of captive breeding programs for the conservation of bighorn sheep. Transactions of the Desert Bighorn Council 36:54-57.

1991; Boyce, W.M., D.A. Jessup and R.K. Clark. Serodiagnostic antibody responses to Psoroptes sp. infestations in bighorn sheep. Journal of Wildlife Diseases 27:10-15.

1991; Boyce, W.M. and R.N. Brown. Antigenic characterization of Psoroptes spp. (Acari: Psoroptidae) mites from different hosts. Journal of Parasitology 77:675-679.

1991; Boyce, W.M., J.A.K. Mazet, J. Mellies, I. Gardner, R.K. Clark and D.A. Jessup. Kinetic ELISA for detection of antibodies to Psoroptes sp. (Acari: Psoroptidae) in bighorn sheep (Ovis canadensis). Journal of Parasitology 77:692-696.

1990; Boyce, W.M., L. Elliott, R.K. Clark and D.A. Jessup. Morphometric analysis of Psoroptes spp. mites from bighorn sheep, mule deer, rabbits and cattle. Journal of Parasitology 76:823-828.

Mesocestoides

2005; Padgett, K.A, and W.M. Boyce. Ants as first intermediate host of Mesocestoides on San Miguel Island, USA. Journal of Helminthology. 79:67-73. pdf

2005; Padgett, K.A, S.A. Nadler, L. Munson, Sacks, B. and W.M. Boyce. Systematics of Mesocestoides (Cestoda: Mesocestoididae):  Evaluation of molecular and morphological variation among isolates. Journal of Parasitology. 91:1435-1443. pdf

2004; Padgett, K.A, and W.M. Boyce. Life history studies on two molecular strains of Mesocestoides (Cestoda: Mesocestoididae): identification of sylvatic hosts and infectivity of immature life stages. Journal of Parasitology. 90:108-113. pdf

2000; Crosbie, P.R., S.A. Nadler, E.G. Platzer, C. Kerner, J. Mariaux and W.M. Boyce. Molecular systematics of Mesocestoides spp. (Cestoda: Mesocestoididae) from domestic dogs (Canis familiaris) and coyotes (Canis latrans). Journal of Parasitology 86:350-357. pdf

1998; Crosbie, P.R., W.M. Boyce, E.G. Platzer, S.A. Nadler and C. Kerner. Diagnostic procedures and treatment of eleven dogs with peritoneal infections caused by Mesocestoides spp. Journal of the American Veterinary Medical Association 213:1578-1583.

Psoroptes

2005; Boyce, W.M. and M. Weisenberger. The rise and fall of psoroptic scabies in bighorn sheep in the San Andres Mountains, New Mexico. Journal of Wildlife Diseases. 41:525—531. pdf

2000; Ramey, R.R., S.T. Kelley, W.M. Boyce and B.D. Farrell. Phylogeny and host specificity of psoroptic mange mites (Acarina:Psoroptidae) as indicated by ITS sequence data. Journal of Medical Entomology 37(6):791-796. pdf

1997; Singer, R.S., D.A. Jessup, I.A. Gardner and W.M. Boyce. Pathogen exposure patterns among sympatric populations of bighorn sheep (Ovis canadensis), mule deer (Odocoileus hemionus), and cattle. Journal of Wildlife Diseases 33:377-382. pdf

1996; Ziccardi, M., W.M. Boyce and I. Gardner. Seroprevalence of Psoroptes sp. mites in free-ranging elk (Cervus elaphus) as determined by kinetic ELISA. Journal of Wildlife Diseases 32:51-56. pdf

1996; Reese, N.E., W.M. Boyce, I.A. Gardner and D.M. Nelson. Fixation affects morphometric characters of Psoroptes cuniculi mites (Acari:Psoroptidae). Journal of Medical Entomology 33:835-838.

1996; Boyce, W.M. and R.L. Zarnke. Antibody responses to Psoroptes sp. mites in Dall sheep (Ovis dalli). Journal of Wildife Diseases 32:711-713

1992; Foreyt, W.J., L.G. Rickard and W.M. Boyce. Psoroptes sp. in two llamas in Washington. Journal of Parasitology 78:153-155.

1992; Boyce, W.M., J.A. Miller, D.A. Jessup and R.K. Clark. Use of ivermectin implants for the treatment of psoroptic scabies in free-ranging bighorn sheep. Journal of Zoo and Wildlife Medicine 23:211-213.

1992; Mazet, J.A.K., W.M. Boyce, J. Mellies, R.K. Clark and D.A. Jessup. Exposure to Psoroptes sp. mites is common among bighorn sheep (Ovis canadensis) populations in California. Journal of Wildlife Diseases 28:542-547. pdf

1991; Boyce, W.M., D.A. Jessup and R.K. Clark. Serodiagnostic antibody responses to Psoroptes sp. infestations in bighorn sheep. Journal of Wildlife Diseases 27:10-15.

1991; Boyce, W.M. and R.N. Brown. Antigenic characterization of Psoroptes spp. (Acari: Psoroptidae) mites from different hosts. Journal of Parasitology 77:675-679.

1991; Boyce, W.M., J.A.K. Mazet, J. Mellies, I. Gardner, R.K. Clark and D.A. Jessup. Kinetic ELISA for detection of antibodies to Psoroptes sp. (Acari: Psoroptidae) in bighorn sheep (Ovis canadensis). Journal of Parasitology 77:692-696.

1990; Boyce, W.M., L. Elliott, R.K. Clark and D.A. Jessup. Morphometric analysis of Psoroptes spp. mites from bighorn sheep, mule deer, rabbits and cattle. Journal of Parasitology 76:823-828.